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In both sexes, the genitalia are adorned with various spines, teeth, scales and bristles, which act to prevent the butterfly from mating with an insect of another species.
A newly emerged butterfly needs to spend some time inflating its wings with hemolymph and letting them dry, during which time it is extremely vulnerable to predators.
Butterflies feed primarily on nectar from flowers. Some also derive nourishment from pollen ,  tree sap, rotting fruit, dung, decaying flesh, and dissolved minerals in wet sand or dirt.
Butterflies are important as pollinators for some species of plants. In general, they do not carry as much pollen load as bees , but they are capable of moving pollen over greater distances.
Adult butterflies consume only liquids, ingested through the proboscis. They sip water from damp patches for hydration and feed on nectar from flowers, from which they obtain sugars for energy, and sodium and other minerals vital for reproduction.
Several species of butterflies need more sodium than that provided by nectar and are attracted by sodium in salt; they sometimes land on people, attracted by the salt in human sweat.
Some butterflies also visit dung and scavenge rotting fruit or carcasses to obtain minerals and nutrients.
In many species, this mud-puddling behaviour is restricted to the males, and studies have suggested that the nutrients collected may be provided as a nuptial gift , along with the spermatophore, during mating.
In hilltopping , males of some species seek hilltops and ridge tops, which they patrol in search for females. Since it usually occurs in species with low population density, it is assumed these landscape points are used as meeting places to find mates.
Butterflies use their antennae to sense the air for wind and scents. The antennae come in various shapes and colours; the hesperiids have a pointed angle or hook to the antennae, while most other families show knobbed antennae.
The antennae are richly covered with sensory organs known as sensillae. A butterfly's sense of taste is coordinated by chemoreceptors on the tarsi , or feet, which work only on contact, and are used to determine whether an egg-laying insect's offspring will be able to feed on a leaf before eggs are laid on it.
Many species show sexual dimorphism in the patterns of UV reflective patches. Many species of butterfly maintain territories and actively chase other species or individuals that may stray into them.
Some species will bask or perch on chosen perches. The flight styles of butterflies are often characteristic and some species have courtship flight displays.
Some species have evolved dark wingbases to help in gathering more heat and this is especially evident in alpine forms.
As in many other insects, the lift generated by butterflies is more than can be accounted for by steady-state, non-transitory aerodynamics.
Studies using Vanessa atalanta in a wind tunnel show that they use a wide variety of aerodynamic mechanisms to generate force.
These include wake capture , vortices at the wing edge, rotational mechanisms and the Weis-Fogh ' clap-and-fling ' mechanism.
Butterflies are able to change from one mode to another rapidly. Butterflies are threatened in their early stages by parasitoids and in all stages by predators, diseases and environmental factors.
Braconid and other parasitic wasps lay their eggs in lepidopteran eggs or larvae and the wasps' parasitoid larvae devour their hosts, usually pupating inside or outside the desiccated husk.
Most wasps are very specific about their host species and some have been used as biological controls of pest butterflies like the large white butterfly.
In order to control it, some pupae that had been parasitised by a chalcid wasp were imported, and natural control was thus regained.
Caterpillars are also affected by a range of bacterial, viral and fungal diseases, and only a small percentage of the butterfly eggs laid ever reach adulthood.
Queen Alexandra's birdwing is the largest butterfly in the world. The species is endangered , and is one of only three 3 insects the other two being butterflies as well to be listed on Appendix I of CITES , making international trade illegal.
Black grass-dart butterfly Ocybadistes knightorum is a butterfly of the family Hesperiidae. It is endemic to New South Wales. It has a very limited distribution in the Boambee area.
Chemical defences are widespread and are mostly based on chemicals of plant origin. In many cases the plants themselves evolved these toxic substances as protection against herbivores.
Butterflies have evolved mechanisms to sequester these plant toxins and use them instead in their own defence. This signal is commonly mimicked by other butterflies, usually only females.
A Batesian mimic imitates another species to enjoy the protection of that species' aposematism. Camouflage is found in many butterflies.
Some like the oakleaf butterfly and autumn leaf are remarkable imitations of leaves. Some female Nymphalid butterflies guard their eggs from parasitoidal wasps.
The Lycaenidae have a false head consisting of eyespots and small tails false antennae to deflect attack from the more vital head region.
These may also cause ambush predators such as spiders to approach from the wrong end, enabling the butterflies to detect attacks promptly.
Auditory defences can also be used, which in the case of the grizzled skipper refers to vibrations generated by the butterfly upon expanding its wings in an attempt to communicate with ant predators.
Many tropical butterflies have seasonal forms for dry and wet seasons. Dark colours in wet-season forms may help to absorb solar radiation. Butterflies without defences such as toxins or mimicry protect themselves through a flight that is more bumpy and unpredictable than in other species.
It is assumed this behavior makes it more difficult for predators to catch them, and is caused by the turbulence created by the small whirlpools formed by the wings during flight.
Butterflies have appeared in art from years ago in ancient Egypt. The butterfly was sometimes depicted with the maw of a jaguar , and some species were considered to be the reincarnations of the souls of dead warriors.
The close association of butterflies with fire and warfare persisted into the Aztec civilisation ; evidence of similar jaguar-butterfly images has been found among the Zapotec and Maya civilisations.
Butterflies are widely used in objects of art and jewellery: mounted in frames, embedded in resin, displayed in bottles, laminated in paper, and used in some mixed media artworks and furnishings.
The caterpillar is seated on a toadstool and is smoking a hookah ; the image can be read as showing either the forelegs of the larva, or as suggesting a face with protruding nose and chin.
Madam Butterfly is a opera by Giacomo Puccini about a romantic young Japanese bride who is deserted by her American officer husband soon after they are married.
It was based on John Luther Long 's short story written in According to Lafcadio Hearn , a butterfly was seen in Japan as the personification of a person's soul; whether they be living, dying, or already dead.
One Japanese superstition says that if a butterfly enters your guest room and perches behind the bamboo screen, the person whom you most love is coming to see you.
Large numbers of butterflies are viewed as bad omens. When Taira no Masakado was secretly preparing for his famous revolt, there appeared in Kyoto so vast a swarm of butterflies that the people were frightened—thinking the apparition to be a portent of coming evil.
A Roman sculpture depicts a butterfly exiting the mouth of a dead man, representing the Roman belief that the soul leaves through the mouth.
Collecting butterflies was once a popular hobby; it has now largely been replaced by photography, recording, and rearing butterflies for release into the wild.
He published the results in the folio sized handbook The Natural History of British Butterflies in Study of the structural coloration of the wing scales of swallowtail butterflies has led to the development of more efficient light-emitting diodes ,  and is inspiring nanotechnology research to produce paints that do not use toxic pigments and the development of new display technologies.
From Wikipedia, the free encyclopedia. For other uses, see Butterfly disambiguation. A group of insects in the order Lepidoptera. Further information: Prehistoric Lepidoptera.
Further information: Glossary of entomology terms and Comparison of butterflies and moths. Further information: Lepidoptera migration , Insect migration , and Animal navigation.
Further information: Defense in insects , Anti-predator adaptation , Mimicry , and Seasonal polyphenism.
Further information: Biomimetics. Bugs Britannica. Chatto and Windus. Evolution of the Insects. Cambridge University Press. Do Butterflies Bite?
Rutgers University Press. Geological Society of America. University of California. Archived from the original on 7 April Retrieved 15 July Bibcode : Geo Retrieved 8 September The Journal of Physical Chemistry.
Ghiradella Photonics Science News. February The Journal of Experimental Biology. The Insects: An Outline of Entomology 5 ed.
Ecological Entomology. Amateur Entomologists' Society. Retrieved 13 September August The Lepidopterists' Society. Archived from the original on 13 May Retrieved 9 September Monarch Lab.
Archived from the original on 6 October Science News. Alfred A. BirdGuides Ltd. Retrieved 22 October Evershed, E. Green, J.
Fryer and W. Transactions of the Entomological Society of London. Bibcode : PNAS Current Biology. Journal of Insect Conservation.
Butterflies and Moths of North America. Retrieved 15 September Insect Ecology: An Ecosystem Approach. Academic Press. Marshall Cavendish. Encyclopedia of Entomology.
Bulletin of Entomological Research. Stanford University Press. UBC Press. Archived from the original on 7 January Retrieved 12 September Archived from the original on 21 April Current Science.
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